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Assessing the origin of Neotropical mountain dung beetle assemblages (Scarabaeidae: Scarabaeinae) : the comparative influence of vertical and horizontal colonization 404

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Assessing the origin of Neotropical mountain dung beetle assemblages (Scarabaeidae: Scarabaeinae) : the comparative influence of vertical and horizontal colonization 71 7 4 1 5 2 8

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