Verster, Anna Johanna Maria; Jansen B.C.
(Pretoria : The Government Printer, 1965)
1. The nominate subspecies of E. granulosus (Batsch, 1786) is redesignated from the type locality, Europe.
2. Five subspecies of E. granulosus are described from South African carnivores: E. g. granulosus, E. g. africanus, ...